I highly doubt you'll be able to find many Perl/MySQL scripts out there for forum software, as everyone's gone the PHP/MySQL route...

I would have serious reservations against WWWThreads, as it hasn't been deved in, forever (especially the perl version), and is definately EOL, I would expect that there be some major security issues as a possibility, and as it isn't developed, they could arise anywhere and you'll have no real method for patching...

Honestly, UBB.C6.7.3 would be my only recommendation in this case; which yes, does do flat file storage (which every perl based forum I've ever used does flat file storage as well).

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