In /templates/default/showflat.tpl you'll see a <DIV> surrounding those three links, around line 194 (in v7.0). You should be able to add a style to that DIV to change the link colors and/or background color.
This is untested, so play with it. Change that <div> to <div class="prevnext"> then add something like this to "Extra Properties" in the Styles Editor:
I'm no CSS expert, but it'd probably be something like this (edit colors accordingly)
.privnext a:link {
color: #222222;
background: none;
}
.privnext a:visited {
color: #222222;
background: none;
}
.privnext a:active {
color: #ff0000;
background: none;
}
.privnext a:hover {
color: #ff0000;
background: none;
}
Do the same for showthreaded.tpl once you got it working...